A number of intermediate precursors of carolic acid in P. charlesii has been studied, the most notable being γ-methyltetronic acid and carlosic acid which appear to be incorporated via separate metabolic pathways.
J. L. Bloomer, F. E. Kappler and G. N. Pandey, J. Chem. Soc., Chem. Commun., 1972, 243 DOI: 10.1039/C39720000243
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