Biogenetic studies indicate that gentisein and related xanthones are derived from a combination of both acetate and shikimate precursors, with the involvement of 2,3′,4,6-tetrahydroxybenzophenone in an oxidative coupling conversion into the xanthone.
P. Gupta and J. R. Lewis, J. Chem. Soc. C, 1971, 629 DOI: 10.1039/J39710000629
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