Four species of Boletus contain a new tetronic acid, variegatic acid, which has been identified as 3,3′,4,4′-tetrahydroxypulvinic acid. The compound is responsible for the blueing reaction in these species.
P. C. Beaumont, R. L. Edwards and G. C. Elsworthy, J. Chem. Soc. C, 1968, 2968 DOI: 10.1039/J39680002968
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