* Corresponding authors

^a Department of Chemical and Biomolecular Engineering, Yonsei University, Seoul 03722, Republic of Korea
E-mail: haam@yonsei.ac.kr

^b Bionanotechnology Research Center Korea Research Institute of Bioscience and Biotechnology (KRIBB), Daejeon 34141, Republic of Korea
E-mail: eklim1112@kribb.re.kr

^c Department of Nanobiotechnology, KRIBB School, University of Science and Technology (UST), 125 Gwahak-ro, Yuseong-gu, Daejeon 34113, Republic of Korea

^d Department of Laboratory Medicine, Gyeongsang National University Changwon Hospital, Changwon 51472, Republic of Korea

^e School of Pharmacy, Sungkyunkwan University, Suwon 16419, Republic of Korea

^f Wide River Institute of Immunology, Seoul National University College of Medicine, Hongchon 25159, Republic of Korea

^g Department of Microbiology and Immunology, Seoul National University College of Medicine, Seoul 03080, Republic of Korea

^h Department of Biomedical Sciences, Seoul National University College of Medicine, Seoul 03080, Republic of Korea

ⁱ Shaperon, Inc., Seoul 06373, Republic of Korea

^j Research Center, HuVet bio, Inc., Seoul 05836, Republic of Korea