Two triterpene-derived aromatic hydrocarbons of the gammacerane-type have been isolated from Messel shale; structural (MS, NMR) and carbon isotopic data indicate that these biological markers are derived from tetrahymanol from protozoa or purple phototrophic bacteria, and from a bacterial hopanoid.
P. Schaeffer, Jean-M. Trendel and P. Albrecht, J. Chem. Soc., Chem. Commun., 1995, 1275 DOI: 10.1039/C39950001275
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